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Cover: Achieve for Ecology: The Economy of Nature 10e with Evolution 4e (1-Term Online), 10th Edition by Rick Relyea; Douglas Emlen; Carl Zimmer
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Achieve for Ecology: The Economy of Nature 10e with Evolution 4e (1-Term Online)

Tenth  Edition|©2026  Rick Relyea; Douglas Emlen; Carl Zimmer

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Authors

Rick Relyea

Rick Relyea  is the William J. Rucker Professor in Fisheries and Wildlife at the University of Missouri–Columbia (Mizzou). He received a BS in environmental forest biology from the SUNY College of Environmental Science and Forestry, an MS in wildlife management from Texas Tech University, and a PhD in ecology and evolution from the University of Michigan. He has authored more than 200 scientific articles and presented research seminars throughout the world. Dr. Relyea was a professor at the University of Pittsburgh for 15 years, where he was named the Chancellor’s Distinguished Researcher and received the Tina and David Bellet Teaching Excellence Award. In 2014, he moved to Rensselaer Polytechnic Institute to direct the Darrin Fresh Water Institute. In 2025, Rick moved to Mizzou to be the founding director of the Johnny Morris Institute of Fisheries, Wetlands, and Aquatic Systems. Rick has a strong interest in high school education, including hosting high school science teachers who conduct research in his laboratory. He is the author of the college textbook Ecology: The Economy of Nature, and co-author of Environmental Science for the AP® Course, which is also published by BFW publishers.


Douglas Emlen

Douglas J. Emlen is a professor at the University of Montana. He is a fellow of the American Academy of Arts and Sciences, a member of the National Academy of Sciences, and a recipient of the U.S. Pres- idential Early Career Award for Scientists and Engineers. In 2014 he was awarded UM’s Distinguished Teaching Award and in 2015 the Carnegie/CASE Professor of the Year Award for the state of Montana. His 2014 book Animal Weapons: The Evolution of Battle won the Phi Beta Kappa Award in Science, and he recently starred in documenta- ries about his work on BBC (Nature’s Wildest Weapons) and NOVA (Extreme Animal Weapons).


Carl Zimmer

Carl Zimmer is one of the country’s leading science writers. A colum- nist for the New York Times, he is the author of 15 books, including She Has Her Mother’s Laugh: The Powers, Perversions, and Poten- tials of Heredity, which The Guardian named the best science book of 2018. Zimmer is professor adjunct at Yale University, where he teaches science writing. Among his many honors, Zimmer has won the Stephen Jay Gould Prize, awarded by the Society for the Study of Evolution, and the National Association of Biology Teachers Distinguished Service Award.


Table of Contents

Headshot of Rick Relyea

Rick Relyea

Rick Relyea  is the William J. Rucker Professor in Fisheries and Wildlife at the University of Missouri–Columbia (Mizzou). He received a BS in environmental forest biology from the SUNY College of Environmental Science and Forestry, an MS in wildlife management from Texas Tech University, and a PhD in ecology and evolution from the University of Michigan. He has authored more than 200 scientific articles and presented research seminars throughout the world. Dr. Relyea was a professor at the University of Pittsburgh for 15 years, where he was named the Chancellor’s Distinguished Researcher and received the Tina and David Bellet Teaching Excellence Award. In 2014, he moved to Rensselaer Polytechnic Institute to direct the Darrin Fresh Water Institute. In 2025, Rick moved to Mizzou to be the founding director of the Johnny Morris Institute of Fisheries, Wetlands, and Aquatic Systems. Rick has a strong interest in high school education, including hosting high school science teachers who conduct research in his laboratory. He is the author of the college textbook Ecology: The Economy of Nature, and co-author of Environmental Science for the AP® Course, which is also published by BFW publishers.


Headshot of Douglas Emlen

Douglas Emlen

Douglas J. Emlen is a professor at the University of Montana. He is a fellow of the American Academy of Arts and Sciences, a member of the National Academy of Sciences, and a recipient of the U.S. Pres- idential Early Career Award for Scientists and Engineers. In 2014 he was awarded UM’s Distinguished Teaching Award and in 2015 the Carnegie/CASE Professor of the Year Award for the state of Montana. His 2014 book Animal Weapons: The Evolution of Battle won the Phi Beta Kappa Award in Science, and he recently starred in documenta- ries about his work on BBC (Nature’s Wildest Weapons) and NOVA (Extreme Animal Weapons).


Headshot of Carl Zimmer

Carl Zimmer

Carl Zimmer is one of the country’s leading science writers. A colum- nist for the New York Times, he is the author of 15 books, including She Has Her Mother’s Laugh: The Powers, Perversions, and Poten- tials of Heredity, which The Guardian named the best science book of 2018. Zimmer is professor adjunct at Yale University, where he teaches science writing. Among his many honors, Zimmer has won the Stephen Jay Gould Prize, awarded by the Society for the Study of Evolution, and the National Association of Biology Teachers Distinguished Service Award.


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