Cover: Achieve for Ecology: The Economy of Nature 10e with Evolution 4e (1-Term Access), 10th Edition by Rick Relyea; Douglas Emlen; Carl Zimmer

Achieve for Ecology: The Economy of Nature 10e with Evolution 4e (1-Term Access)

Tenth  Edition©2026  Rick Relyea; Douglas Emlen; Carl Zimmer

About

Build on your biology foundation with a course that brings ecology and evolution together. This Achieve edition combines Relyea’s Ecology: The Economy of Nature and Emlen/Zimmer’s Evolution: Making Sense of Life into one seamless, easy-to-use Achieve platform. Explore how life evolves and ecosystems function—through real-world examples, smart study tools, and interactive assignments designed to help you succeed.

Table of Contents

Relyea, Economy of Nature 10e

1 An Introduction to Ecology

Part I: Climates, Climate Change, and Biomes

2 Global Climates

3 Global Climate Change

4 Terrestrial and Aquatic Biomes

Part II: Adaptations to Environments

5 Evolutionary Ecology

6 Adaptations to Aquatic Environments

7 Adaptations to Terrestrial Environments

8 Adaptations to Variable Environments

Part III: Life Histories, Reproductive Strategies, and Social Behaviors

9 Life Histories

10 Reproductive Strategies

11 Social Behaviors

Part IV: Populations

12 Population Distributions

13 Population Growth and Regulation

14 Population Dynamics over Time and Space

Part V: Species Interactions

15 Predation and Herbivory

16 Parasitism and Infectious Diseases

17 Competition

18 Mutualism

Part VI: Communities and Ecosystems

19 Community Structure: Biodiversity and Food Webs

20 Community Succession

21 Energy Flow in Ecosystems

22 Nutrient Cycling in Ecosystems

Part VII: Global Ecology

23 Landscape Ecology and Global Biodiversity

24 Conservation of Global Biodiversity

Emlen, Evolution 4e

1 The Whale and the Virus: How Scientists Study Evolution

2 From Natural Philosophy to Darwin: A Brief History of Evolutionary Ideas 

3 What the Rocks Say: How Geology and Paleontology Reveal the History of Life 

4 The Tree of Life: How Biologists Use Phylogeny to Reconstruct the Deep Past 

5 Raw Material: Heritable Variation Among Individuals 

6 The Ways of Change: Drift and Selection

7 Beyond Alleles: Quantitative Genetics and the Evolution of Phenotypes

8 The History in Our Genes 

9 From Genes to Traits: The Evolution of Genetic Networks and Development

10 Natural Selection: Empirical Studies in the Wild

11 Sex: Causes and Consequences 

12 After Conception: The Evolution of Life History and Parental Care 

13 The Origin of Species 

14 Macroevolution: The Long Run 

15 Intimate Partnerships: How Species Adapt to Each Other 

16 Brains and Behavior 

17 Human Evolution: A New Kind of Ape 

18 Evolutionary Medicine 

Rick Relyea

Rick Relyea  is the William J. Rucker Professor in Fisheries and Wildlife at the University of Missouri–Columbia (Mizzou). He received a BS in environmental forest biology from the SUNY College of Environmental Science and Forestry, an MS in wildlife management from Texas Tech University, and a PhD in ecology and evolution from the University of Michigan. He has authored more than 200 scientific articles and presented research seminars throughout the world. Dr. Relyea was a professor at the University of Pittsburgh for 15 years, where he was named the Chancellor’s Distinguished Researcher and received the Tina and David Bellet Teaching Excellence Award. In 2014, he moved to Rensselaer Polytechnic Institute to direct the Darrin Fresh Water Institute. In 2025, Rick moved to Mizzou to be the founding director of the Johnny Morris Institute of Fisheries, Wetlands, and Aquatic Systems. Rick has a strong interest in high school education, including hosting high school science teachers who conduct research in his laboratory. He is the author of the college textbook Ecology: The Economy of Nature, and co-author of Environmental Science for the AP® Course, which is also published by BFW publishers.

Douglas Emlen

Douglas J. Emlen is a professor at the University of Montana. He is a fellow of the American Academy of Arts and Sciences, a member of the National Academy of Sciences, and a recipient of the U.S. Pres- idential Early Career Award for Scientists and Engineers. In 2014 he was awarded UM’s Distinguished Teaching Award and in 2015 the Carnegie/CASE Professor of the Year Award for the state of Montana. His 2014 book Animal Weapons: The Evolution of Battle won the Phi Beta Kappa Award in Science, and he recently starred in documenta- ries about his work on BBC (Nature’s Wildest Weapons) and NOVA (Extreme Animal Weapons).

Carl Zimmer

Carl Zimmer is one of the country’s leading science writers. A colum- nist for the New York Times, he is the author of 15 books, including She Has Her Mother’s Laugh: The Powers, Perversions, and Poten- tials of Heredity, which The Guardian named the best science book of 2018. Zimmer is professor adjunct at Yale University, where he teaches science writing. Among his many honors, Zimmer has won the Stephen Jay Gould Prize, awarded by the Society for the Study of Evolution, and the National Association of Biology Teachers Distinguished Service Award.